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h2{font-size:clamp(45px,12vw,65px);line-height:.96;letter-spacing:-.035em}.section-intro{font-size:14px;max-width:40ch;margin-top:24px}.layer-subtitle{font-size:11px;margin-bottom:25px;max-width:32ch}.map-panel{grid-template-columns:1fr;padding:28px 20px;gap:30px;margin-inline:-8px}.map-image{max-width:360px}.map-figure figcaption{font-size:10px;text-align:left;max-width:38ch;margin-top:18px}.hotspot{left:82%;min-width:44px;min-height:44px;font-size:11px}.hotspot:after{width:16px}.layer-controls>.eyebrow{font-size:9px;margin-bottom:15px}.layer-selector{min-height:86px;gap:14px;padding:18px 10px 18px 14px}.layer-name{font-size:29px}.layer-theme{font-size:12px}.layer-number{font-size:11px}.layer-dot{width:10px;height:10px}.selector-arrow{font-size:20px}.layer-detail{min-height:0;border-left:0;border-top:1px solid var(--ink-rule);padding:30px 0 10px}.layer-detail h3{font-size:42px;max-width:15ch;margin-bottom:24px}.layer-detail .detail-kicker{font-size:10px}.layer-detail .original-copy{font-size:28px!important}.layer-detail>p:not(.eyebrow):not(.literal-label){font-size:15px}.gloss-block>p:last-child{font-size:14px}.overview-names{font-size:13px;gap:9px}.map-footnote{font-size:11px;margin-top:20px}.static-layers{grid-template-columns:1fr}.atlas-section{display:flex;flex-direction:column;gap:44px}.atlas-visual{position:static;width:100%}.atlas-visual figure img{height:440px;object-position:center 45%}.atlas-visual figcaption{font-size:8px;margin-top:0;padding-top:13px}.atlas-visual>.button{margin-top:22px;font-size:12px}.atlas-content{width:100%}.atlas-content h2{font-size:54px;line-height:.95}.atlas-content .section-label{margin-bottom:22px}.archive-intro{font-size:14px;margin-top:25px}.search-block{margin-top:27px}.search-block label{font-size:12px}.search-row{gap:9px}#topic-search{font-size:12px;padding-inline:0}.clear-search{font-size:10px;min-width:69px}.topics{margin-top:20px}.topic>summary{min-height:87px;gap:12px;padding-block:20px}.topic-number{min-width:18px;font-size:9px}.topic-name{font-size:28px;max-width:24ch}.topic-plus{font-size:25px;min-width:16px;text-align:center}.topic-body{padding:2px 0 27px}.topic-body dt{font-size:11px}.topic-body dd{font-size:14px}.source-label{font-size:8px}.landscape-break{height:320px}.landscape-break>figcaption{bottom:25px;left:22px;right:22px;gap:20px;font-size:8px;letter-spacing:.1em}.full-table h2{font-size:52px;line-height:1;align-items:flex-start}.full-table h2>span{font-size:30px;padding-top:8px}.full-table>summary{min-height:70px}.table-intro{display:block;margin-top:24px;padding-top:20px}.table-intro p{font-size:11px}.table-intro p+p{margin-top:10px}.table-scroll-hint{font-size:11px}.table-scroll table{min-width:1100px}.table-scroll th,.table-scroll td{padding-top:20px}.site-footer{padding:48px 0 24px}.footer-top{grid-template-columns:1fr;gap:28px}.footer-wordmark{font-size:42px}.footer-by{font-size:9px}.footer-paths{font-size:12px;gap:3px}.footer-paths>a{max-width:none}.brand-plaque{justify-self:start;margin-top:6px;padding:3px 12px}.brand-plaque img{width:135px}.footer-bottom{display:block;margin-top:36px;padding-top:24px;font-size:10px}.footer-bottom>span{display:block;text-align:left;margin-top:20px;font-size:9px}.footer-bottom>span br{display:none}}
@media(prefers-reduced-motion:reduce){*,*::before,*::after{scroll-behavior:auto!important;transition:none!important;animation:none!important}}

/* Main-site navigation is separate from the four study anchors. */
.nav-links .nav-home{display:none}.footer-paths>.main-page-link{color:var(--ink);margin-top:12px;max-width:260px;padding-inline:18px}.footer-paths>.main-page-link:hover{text-decoration:none}@media(max-width:1099px){.nav-links .nav-home{display:flex;padding:9px 12px;gap:12px;min-height:44px;color:var(--ink);font-size:11px}}@media(max-width:767px){.nav-links .nav-home{margin-top:12px;padding:13px 16px;font-size:12px}}
/* Keep keyboard focus visible when a control's ink text sits on dark media. */
.hero .button:focus-visible,.site-footer .button:focus-visible,.map-panel button:focus-visible{outline:2px solid var(--ivory);outline-offset:5px;box-shadow:0 0 0 3px var(--ink)}

/* Study materials: one available pair and one honest processing state. */
.materials-section{background:var(--ink);color:var(--ivory)}.materials-section .label-rule{background:var(--ink-rule)}.materials-section .section-label>span:last-child{color:var(--muted-ink)}.materials-heading{display:grid;grid-template-columns:minmax(0,1.7fr) minmax(0,.65fr);gap:64px;align-items:end;margin:30px 0 64px}.materials-heading h2{font-size:clamp(3rem,5.8vw,6rem);max-width:19ch}.materials-heading>p{color:var(--muted-ink);font-size:14px;max-width:34ch;padding-bottom:4px}.module-entry{display:grid;grid-template-columns:minmax(0,.95fr) minmax(0,1.05fr);gap:clamp(32px,5vw,80px);padding-bottom:64px}.module-cover{min-width:0}.module-cover>a{display:block;aspect-ratio:900/502;background:#242925}.module-cover img{width:100%;height:100%;object-fit:contain}.module-cover figcaption{font-size:10px;letter-spacing:.12em;color:var(--muted-ink);margin-top:16px}.module-content{min-width:0}.module-kicker{display:flex;justify-content:space-between;align-items:center;gap:16px;margin-bottom:20px}.module-status{font-size:11px;letter-spacing:.09em;text-transform:uppercase;color:var(--coral)}.module-content h3{font-size:clamp(42px,4.8vw,68px);line-height:1;letter-spacing:-.025em;max-width:16ch}.module-document-links{display:flex;align-items:center;flex-wrap:wrap;gap:12px 28px;margin-top:28px}.module-document-links .button{gap:28px}.module-document-links .button>span[aria-hidden]{font-size:20px;line-height:1}.module-download{display:inline-flex;align-items:center;justify-content:space-between;gap:22px;min-height:44px;font-size:12px;text-decoration:none}.module-download:hover{text-decoration:underline}.module-download>span{font-size:20px}.module-podcast{border-top:1px solid var(--ink-rule);padding-top:24px;margin-top:36px}.podcast-heading{display:flex;align-items:baseline;justify-content:space-between;gap:16px;margin-bottom:16px}.podcast-heading h4{font:400 30px/1.1 var(--serif);margin:0}.podcast-heading>p{color:var(--muted-ink);font-size:11px;white-space:nowrap}.module-podcast audio{display:block;width:100%;min-width:0;height:54px;color-scheme:light}.podcast-download{margin-top:12px;color:var(--muted-ink)}.module-pending{display:grid;grid-template-columns:minmax(0,.5fr) minmax(0,1.2fr) minmax(0,1fr);gap:40px;border-top:1px solid var(--ink-rule);padding-top:40px;padding-bottom:40px}.module-pending .eyebrow{color:var(--muted-ink)}.module-pending .module-status{margin-top:12px}.module-pending h3{font-size:clamp(34px,3.5vw,50px);line-height:1.05;letter-spacing:-.02em}.processing-notice{font-size:16px;line-height:1.6;max-width:28ch;align-self:center;color:var(--muted-ink)}.materials-coming{display:flex;align-items:baseline;justify-content:space-between;gap:32px;border-top:1px solid var(--ink-rule);padding-top:28px}.materials-coming h3{font-size:32px;line-height:1.1}.materials-coming>p{color:var(--muted-ink);font-size:13px;max-width:46ch}.materials-section .button:focus-visible,.materials-section audio:focus-visible{outline:2px solid var(--ivory);outline-offset:5px;box-shadow:0 0 0 3px var(--ink)}
@media(min-width:768px) and (max-width:1023px){.site-header nav{position:static}.js .nav-links{position:absolute;left:var(--gutter);right:var(--gutter);top:105px;z-index:4;display:flex;flex-direction:column;align-items:stretch;gap:4px;background:var(--ink);padding:18px 24px 24px;border-block:1px solid var(--ink-rule)}.js .nav-links a{font-size:13px;min-height:48px}.nav-links .nav-home{margin-top:12px;padding:13px 16px;font-size:12px}}
@media(max-width:1099px){.materials-heading{grid-template-columns:1.4fr 1fr;gap:32px;margin-bottom:44px}.module-entry{gap:32px}.module-pending{grid-template-columns:minmax(0,.7fr) minmax(0,1.3fr);gap:24px}.module-pending .processing-notice{grid-column:2;max-width:36ch}.module-document-links{gap:8px 20px}}
@media(max-width:767px){.materials-heading{display:block;margin-top:26px;margin-bottom:40px}.materials-heading h2{font-size:clamp(43px,11.5vw,58px);line-height:.98;max-width:17ch}.materials-heading>p{margin-top:24px;max-width:40ch}.module-entry{grid-template-columns:1fr;gap:32px;padding-bottom:40px}.module-cover figcaption{font-size:9px;margin-top:13px}.module-content h3{font-size:46px;max-width:15ch}.module-kicker{margin-bottom:16px}.module-document-links{margin-top:24px;gap:8px 20px}.module-document-links .button{padding-inline:18px;font-size:12px}.module-podcast{margin-top:28px}.podcast-heading{gap:12px}.podcast-heading h4{font-size:28px}.podcast-heading>p{font-size:10px}.module-pending{display:block;padding-block:30px}.module-pending h3{font-size:38px;margin-top:24px}.module-pending .processing-notice{margin-top:20px;max-width:32ch}.materials-coming{display:block;padding-top:24px}.materials-coming h3{font-size:32px}.materials-coming>p{margin-top:16px}}
@media(max-width:1023px){html:not(.js) .site-header{flex-wrap:wrap;padding-block:20px}html:not(.js) .site-header nav{margin-left:0;width:100%}html:not(.js) .nav-links{flex-wrap:wrap;gap:8px 20px}}

/* Registration unlocks server-protected materials; no personal data in browser storage. */
.module-gate{margin-top:28px}.module-gate>p{font-size:14px;color:var(--muted-ink);max-width:38ch;margin-bottom:22px}.module-gate .button{gap:30px}.module-gate .button>span{font-size:20px}.registration-section{display:grid;grid-template-columns:minmax(0,.85fr) minmax(0,1.15fr);gap:clamp(40px,6vw,96px);border-top:1px solid var(--ink-rule);padding:48px 0 64px}.registration-intro>.eyebrow{color:var(--muted-ink);margin-bottom:24px}.registration-intro h2{font-size:clamp(46px,5vw,74px);line-height:.98}.registration-intro>p:not(.eyebrow){max-width:39ch;font-size:14px;color:var(--muted-ink);margin-top:24px}.registration-intro>.registration-privacy{font-size:11px!important;max-width:43ch}.registration-content{min-width:0}.registration-connection{font-size:12px;color:var(--muted-ink);margin-bottom:20px}.registration-retry{display:block;border:0;border-bottom:1px solid var(--ivory);background:none;color:var(--ivory);min-height:44px;font-size:13px;margin-bottom:24px;padding:8px 0}#registration-form fieldset{margin:0;padding:0;border:0;min-width:0}#registration-form fieldset:disabled{opacity:.55}.registration-field{margin-bottom:24px}.registration-field label{display:block;font-size:13px;font-weight:500;margin-bottom:9px}.registration-field label>span{font-weight:400;color:var(--muted-ink)}.registration-field input,.registration-field textarea{display:block;width:100%;min-width:0;min-height:48px;border:1px solid #768071;border-radius:0;background:var(--raised);color:var(--ivory);padding:12px 14px;font:400 16px/1.5 var(--sans)}.registration-field textarea{resize:vertical;min-height:104px}.registration-field input[aria-invalid='true'],.registration-field textarea[aria-invalid='true']{border-color:#ffa390}.registration-help{font-size:11px;color:var(--muted-ink);margin-top:8px}.registration-field-error{font-size:12px;color:#ffa390;margin-top:8px}.registration-consent{margin:4px 0 22px}.registration-consent label{display:flex;align-items:flex-start;gap:13px;padding:12px 0;cursor:pointer;font-size:12px;line-height:1.7}.registration-consent input{flex-shrink:0;width:20px;height:20px;margin:1px 0 0;accent-color:var(--coral)}.registration-errors{border:1px solid #ffa390;color:#ffa390;padding:16px;margin-bottom:24px;font-size:13px}.registration-errors>p{font-weight:600}.registration-errors ul{margin:8px 0 0;padding-left:20px}.registration-errors a{display:inline-flex;align-items:center;min-height:44px}.registration-completed{border-top:2px solid var(--coral);padding-top:24px}.registration-confirmation{font:400 36px/1.1 var(--serif);max-width:24ch}.registration-completed>p+p{font-size:13px;color:var(--muted-ink);margin-top:20px;max-width:48ch}.registration-completed>.module-download{margin-top:24px}.registration-nojs{font-size:13px;color:var(--muted-ink);margin-top:20px}.registration-section input:focus-visible,.registration-section textarea:focus-visible,.registration-section button:focus-visible,.registration-section [tabindex='-1']:focus{outline:2px solid var(--ivory);outline-offset:5px;box-shadow:0 0 0 3px var(--ink)}.registration-section .button:disabled{cursor:wait;opacity:.65}.site-footer .admin-link{font-size:10px;color:var(--muted-ink)}
@media(max-width:767px){.registration-section{grid-template-columns:1fr;gap:32px;padding-block:36px 48px}.registration-intro h2{font-size:50px}.registration-intro>p:not(.eyebrow){max-width:40ch}.registration-intro>.eyebrow{margin-bottom:20px}.registration-field{margin-bottom:22px}.registration-consent label{font-size:12px}.registration-confirmation{font-size:32px}#registration-submit{font-size:12px;padding-inline:18px;gap:20px}.module-gate .button{font-size:12px;padding-inline:18px;gap:22px}}

/* Complete source tables share a harmonized four-color family, with ink text. */
.atlas-filters{display:flex;flex-wrap:wrap;gap:8px;margin-top:24px}.atlas-filters button{display:inline-flex;align-items:center;justify-content:space-between;gap:14px;min-height:44px;border:1px solid var(--rule);background:transparent;color:var(--ink);padding:10px 12px;font:500 11px/1.4 var(--sans);text-align:left}.atlas-filters button>span{font-size:10px}.atlas-filters button[aria-pressed='true']{background:var(--ink);border-color:var(--ink);color:var(--ivory)}.topic-table-link{display:inline-flex;align-items:center;min-height:44px;font-size:12px;margin-top:18px}.layer-atlas-link{display:inline-flex;align-items:center;min-height:44px;font-size:12px;line-height:1.5;margin-top:18px;color:var(--ivory)}.selected-record-count{font-size:11px!important;margin-top:20px}.selected-table-context{grid-column:1/-1;border-top:1px solid var(--ink-rule);padding-top:30px}.selected-table-context>.source-label{color:var(--muted-ink);margin-bottom:14px}.selected-table-context>h3{font-size:36px;line-height:1.1}.selected-context-title{font-size:12px;color:var(--muted-ink);margin-top:16px}.selected-context-grid{display:grid;grid-template-columns:1fr 1fr;gap:24px 40px;margin-top:24px}.selected-context-grid>p{font-size:14px;line-height:1.7;color:var(--ivory)}.map-panel:has(.layer-detail[data-layer]){align-items:start}.module-free-note{font-size:13px;color:var(--muted-ink);max-width:42ch;margin-top:22px}.full-source-intro{max-width:76ch;margin-block:36px 28px}.full-source-intro>.source-label{margin-bottom:16px}.full-source-intro>p:last-child{font-size:17px;line-height:1.7}.tables-attribution{font-size:11px;letter-spacing:.08em;text-transform:uppercase}.table-independent-note{font-size:12px;color:var(--muted);margin-top:12px;max-width:75ch}.layer-table{--table-bg:#f3e8bd;--table-head:#e8d48b;--table-alt:#eee0a7;--table-edge:#9f7922;background:var(--table-bg);color:var(--ink);border-top:4px solid var(--table-edge);padding:28px;margin-top:36px}.layer-table[data-source-layer='mesodermo-antiguo']{--table-bg:#f4dfc8;--table-head:#e7bb91;--table-alt:#eed1b0;--table-edge:#a66029}.layer-table[data-source-layer='mesodermo-nuevo']{--table-bg:#f2d6cf;--table-head:#dfa99d;--table-alt:#eac3b9;--table-edge:#a94d3e}.layer-table[data-source-layer='ectodermo']{--table-bg:#dce8ed;--table-head:#b5d1df;--table-alt:#ccdee7;--table-edge:#326d8b}.layer-table>summary{display:flex;align-items:flex-start;justify-content:space-between;gap:24px;list-style:none;cursor:pointer;padding:20px;background:var(--table-head);margin:-4px -4px 0}.layer-table>summary::-webkit-details-marker{display:none}.layer-table>summary h3{font:500 clamp(17px,1.6vw,24px)/1.4 var(--sans);letter-spacing:.01em;max-width:65ch}.layer-table-toggle{font:400 30px/1 var(--serif);flex-shrink:0}.layer-table[open]>.layer-table-toggle{transform:rotate(45deg)}.layer-table[open]>summary>.layer-table-toggle{transform:rotate(45deg)}.table-context{display:grid;grid-template-columns:1fr 1fr;gap:24px 32px;padding-top:28px}.table-context>p{font-size:15px;line-height:1.7}.layer-table .table-scroll{margin-top:18px}.layer-table .table-scroll table{font-size:16px}.layer-table .table-scroll caption{color:var(--ink);font-size:11px}.layer-table .table-scroll thead{background:var(--table-head)}.layer-table .table-scroll thead th{border-top-color:var(--table-edge);padding:18px 14px;font-size:12px}.layer-table .table-scroll tbody tr:nth-child(even){background:var(--table-alt)}.layer-table .table-scroll tbody th,.layer-table .table-scroll tbody td{padding:20px 14px;border-top-color:var(--table-edge)}.layer-table .table-scroll tbody th{font:400 24px/1.2 var(--serif)}.layer-table .table-scroll-hint{color:var(--ink);font-size:11px}.hotspot{left:90%;top:12.5%}.hotspot[data-layer='mesodermo-antiguo']{top:37%}.hotspot[data-layer='mesodermo-nuevo']{top:62%}.hotspot[data-layer='ectodermo']{top:87%}
@media(max-width:767px){.selected-context-grid{grid-template-columns:1fr;gap:20px}.selected-context-grid>p{font-size:14px}.selected-table-context>h3{font-size:32px}.selected-context-title{font-size:11px}.atlas-filters button{font-size:11px;gap:10px}.full-source-intro>p:last-child{font-size:16px}.layer-table{padding:18px;margin-inline:-4px;margin-top:28px}.layer-table>summary{padding:16px;gap:16px}.layer-table>summary h3{font-size:16px}.table-context{grid-template-columns:1fr;gap:20px}.table-context>p{font-size:14px}.hotspot{left:90%}.registration-intro h2{font-size:44px}.registration-confirmation{font-size:30px}}

/* Editorial controls: flat, named, counted and coordinated with the source tables. */
.atlas-filters{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:8px 10px;padding-top:4px}.atlas-filters button{--filter-tint:#ece9df;--filter-edge:#8a8e83;appearance:none;-webkit-appearance:none;box-shadow:none;border-radius:0;border:1px solid var(--filter-edge);background:var(--filter-tint);color:var(--ink);padding:12px 14px;min-height:48px;gap:10px;font-size:12px;line-height:1.45;transition:background-color 160ms ease-out,border-color 160ms ease-out}.atlas-filters button[data-atlas-filter='all']{grid-column:1/-1}.atlas-filters button[data-atlas-filter='endodermo']{--filter-tint:#f3e8bd;--filter-edge:#9f7922}.atlas-filters button[data-atlas-filter='mesodermo-antiguo']{--filter-tint:#f4dfc8;--filter-edge:#a66029}.atlas-filters button[data-atlas-filter='mesodermo-nuevo']{--filter-tint:#f2d6cf;--filter-edge:#a94d3e}.atlas-filters button[data-atlas-filter='ectodermo']{--filter-tint:#dce8ed;--filter-edge:#326d8b}.atlas-filters button>span{font-size:11px;flex-shrink:0;min-width:16px;text-align:right;font-variant-numeric:tabular-nums}.atlas-filters button:hover{border-color:var(--ink)}.atlas-filters button[aria-pressed='true']{background:var(--ink);border:2px solid var(--ink);padding:11px 13px;color:var(--ivory);font-weight:600}.atlas-filters button:focus-visible{outline:2px solid var(--ink);outline-offset:4px}.footer-share{display:grid;grid-template-columns:minmax(0,1fr) auto;align-items:center;gap:48px;border-top:1px solid var(--ink-rule);padding-top:40px;margin-top:48px}.share-copy>.eyebrow{font-size:9px;color:var(--muted-ink);margin-bottom:20px}.share-copy h3{font-size:clamp(36px,4vw,54px);line-height:1.05}.share-url{display:inline-flex;align-items:center;min-height:44px;margin-top:18px;font-size:14px;overflow-wrap:anywhere}.share-copy>p:last-child{font-size:12px;color:var(--muted-ink);margin-top:12px;max-width:48ch}.share-qr{min-width:0}.share-qr>a{display:block;width:160px;background:#fff}.share-qr img{width:160px;height:160px}.share-qr figcaption{margin-top:12px}.share-qr figcaption>a{display:inline-flex;align-items:center;justify-content:space-between;gap:16px;min-height:44px;font-size:11px}.share-qr figcaption span{font-size:18px}.footer-share+.footer-bottom{margin-top:36px}
@media(max-width:767px){.atlas-filters{gap:8px}.atlas-filters button{font-size:11px;gap:8px;padding:12px 11px}.atlas-filters button[aria-pressed='true']{padding:11px 10px}.footer-share{grid-template-columns:1fr;gap:28px;padding-top:32px;margin-top:36px}.share-copy h3{font-size:38px}.share-url{font-size:12px}.share-qr>a,.share-qr img{width:160px;height:160px}.share-qr{justify-self:start}}

/* Closed-body overview and selected anatomical/cellular sprite crops share one stable stage. */
.map-display{height:600px;max-width:520px;display:flex;align-items:center;justify-content:center;--row-index:0}.map-display>.map-closed-image{width:100%;height:100%;aspect-ratio:auto;object-fit:contain}.map-selected-image{width:min(100%,420px);display:grid;gap:12px;align-content:center}.map-state-crop{position:relative;width:100%;aspect-ratio:16/11;overflow:hidden}.map-state-crop>img{position:absolute;left:0;top:0;width:220%;max-width:none;height:auto;aspect-ratio:1100/1375;transform:translateY(calc(var(--row-index)*-25%));pointer-events:none}.map-cell-crop>img{left:-90%}.map-display[data-map-layer='endodermo']{--row-index:0}.map-display[data-map-layer='mesodermo-antiguo']{--row-index:1}.map-display[data-map-layer='mesodermo-nuevo']{--row-index:2}.map-display[data-map-layer='ectodermo']{--row-index:3}
@media(max-width:767px){.map-display{height:460px;max-width:360px}.map-selected-image{gap:12px;width:min(100%,314px)}}
.map-display[data-map-layer='mesodermo-nuevo'] .map-body-crop{aspect-ratio:16/9}

.module-entry+.module-entry{border-top:1px solid var(--ink-rule);padding-top:48px}